From: Subject: IDENTIFICATION, BIOLOGY AND SAMPLING OF COTTON INSECTS Date: Wed, 26 Sep 2007 14:28:21 -0500 MIME-Version: 1.0 Content-Type: multipart/related; type="text/html"; boundary="----=_NextPart_000_0000_01C80049.7D04FA70" X-MimeOLE: Produced By Microsoft MimeOLE V6.00.2900.3138 This is a multi-part message in MIME format. ------=_NextPart_000_0000_01C80049.7D04FA70 Content-Type: text/html; charset="Windows-1252" Content-Transfer-Encoding: quoted-printable Content-Location: http://insects.tamu.edu/extension/bulletins/b-933.html IDENTIFICATION, BIOLOGY AND SAMPLING OF COTTON = INSECTS

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B-933

IDENTIFICATION, BIOLOGY AND SAMPLING OF COTTON INSECTS

G.T. Bohmfalk, R.E. Frisbie, W.L. Sterling, R.B. Metzer and A.E. = Knutson*=20

* Respectively, former Extension assistant, integrated pest = management=20 coordinator, professor of entomology, Extension cotton specialist and = Extension=20 agent-entomology (PM), The Texas A&M University System.

INT= RODUCTION

IDENTI= FICATION,=20 BIOLOGY, EVIDENCE OF INFESTATION AND NATURE OF DAMAGE

= KEY=20 PESTS

Boll=20 weevil
Cotton=20 fleahopper=20
Bollworm=20 and tobacco budworm

OCCASIONAL=20 PESTS=20

Beet=20 armyworm
Brown=20 cotton leafworm
Cabbage=20 looper
Cotton=20 aphid=20
Cotton=20 leafworm=20
Cotton=20 leafperforator=20
Cotton=20 square borer=20
Cotton=20 stainer=20
= Cutworms=20
Fall=20 armyworm=20
Grasshoppers=20
Tarnished=20 plant bug=20
Omnivorous=20 leafroller
Pink=20 bollworm=20
Saltmarsh=20 caterpillar=20
Spider=20 mites=20
Stink=20 Bugs=20
Th= rips=20
Whiteflies=20
Yellow-striped=20 armyworm

PREDACEOUS=20 INSECTS AND SPIDERS

<= STRONG>Insects=20
Minute=20 pirate bugs=20
B= igeyed=20 bugs=20
Ants= =20
Lady=20 beetles=20
Cotton=20 fleahoppers=20
Green=20 lacewings
Damsel=20 bugs
= Assassin=20 bugs=20
<= STRONG>Spiders=20
Striped=20 Iynx=20
Cel= er crab=20 spider=20
Winter=20 spider=20
Star-bellied=20 orb weaver=20
Gr= ey=20 dotted spider=20
Black=20 and white jumping spider=20
Long=20 jawed orb weaver=20
Ridgefaced=20 crab spider
Use=20 of Predators

INSECT=20 PARASITISM OF COTTON PESTS

THE=20 COTTON PLANT

= CROP=20 MONITORING (SCOUTING)

MONITORING=20 COTTON FRUITING RATE

MANAGEMENT=20 DECISIONS

OTHER=20 INSECT PEST MANAGEMENT PRACTICES

INSECT=20 IDENTIFICATION AND DAMAGE-DIAGNOSTIC KEY

ADDITIONAL=20 BENEFICIAL INSECTS

INTRODUCTION

R.E. Frisbie=20

The management of insect pests is an integral part of an economic = production=20 system. It increases producers' profits and reduces the amount of = environmental=20 contamination from pesticides. Integrated pest management (IPM) is the=20 integration of all practical pest control methods in as compatible = manner as=20 possible to maintain pest populations below the economic threshold. In a = cotton=20 production system, IPM includes combining cultural = practices--appropriate=20 variety selection, land preparation, planting dates and early stalk = destruction=20 -- frequent field scouting; biological control through conservation of = natural=20 predators, parasites and pathogens; and selective use of insecticides to = keep=20 insect and mite populations below economically damaging levels. This = cotton pest=20 management system is designed to keep inputs at a minimum while = maintaining or=20 increasing production for maximum profit.=20

This publication covers the identification, biology, life cycle, = evidence of=20 infestation and nature of damage of pests that attack cotton. A special = section=20 deals with recognizing predaceous insects and spiders. Crop monitoring=20 (scouting) is stressed as a method of determining the abundance of pests = and=20 beneficial insect and spider species. The economic threshold (action = level) is=20 related to scouting procedures. Easy-to-use diagnostic keys to aid in=20 recognizing insect damage and identifying eggs and larvae are included.=20

This publication is a guide to a better understanding of the cotton = insect,=20 spider and mite complex. No reference is made to specific control = procedures=20 within geographic areas of the state. The Management of Cotton = Insects=20 series is designed to provide specific management recommendations by = geographic=20 production area. Because of changing needs and constant shifts in = research=20 technology, it is recommended that information on scouting techniques, = economic=20 thresholds, appropriate pesticide use and specific cultural practices be = obtained from the following:=20

B-1210 Management of Cotton Insects in the Lower Rio Grande = Valley of=20 Texas

B-1204 Management of Cotton Insects in South and East Texas=20

B-1209 Management of Cotton Insects in the High Plains, Rolling = Plains=20 and Trans-Pecos Areas of Texas=20

B-1205 Management of Cotton Insects in the Texas Blacklands =

IDENTIFICATION, BIOLOGY, EVIDENCE OF
INFESTATION AND NATURE OF=20 DAMAGE

KEY PESTS
G.T. Bohmfalk

Key pests are serious, perennially occurring, persistent pests that = dominate=20 control practices because in the absence of control their populations = may cause=20 severe economic damage. The following key pests vary in their severity = and=20 prominence by geographical location. None are key pests on a statewide = basis nor=20 in every production year.

Boll weevil (Anthonomous grandis = grandis=20 Boheman)=20

Identification. The adult boll weevil is a brown to = grayish-brown=20 beetle. The body is covered with short, fine hair, giving it a fuzzy = appearance.=20 There is considerable variation in size from slightly more than 1/8 inch = to=20 almost =BD inch in length. The boll weevil's snout is approximately half = as long=20 as its body. It is slightly curved and has chewing mouthparts on the = end.=20 Immature stages are found inside squares and bolls. The boll weevil egg = is=20 seldom seen since it is deposited inside a square or boll. The larva is = a small,=20 legless grub with a brownish head and chewing mouthparts. This grub = varies in=20 size from very small to =BD inch in length. The pupal or "resting" stage = of the=20 boll weevil is 3/8 to =BD inch long and cream colored with eyes and an = obvious=20 snout.=20

Biology. The adult boll weevil spends the winter in = hibernation,=20 called "diapause," without food and returns to cotton in the early = spring the=20 following year. Overwintering quarters usually consist of fence rows,=20 broadleaved plant litter along creek bottoms, ditch banks and other = protected,=20 wooded areas near cotton fields. In the spring overwintered boll weevil = adults=20 concentrate in early planted fields nearest overwintering habitat where = cotton=20 is squaring. Adult boll weevils feed on tender growth in plant terminals = if the=20 young cotton does not have squares. In the early season, boll weevils = colonize=20 localized spots and do not generally invade the entire field.=20

The boll weevil is a pollen feeder; its survival is diminished = without=20 squaring cotton, although adult boll weevils emerging from overwintering = quarters may subsist on other plants for short periods (e.g., an average = of 18=20 days on yellow woolywhite in the Rolling Plains area). After adult = weevils feed=20 on cotton for 3 to 7 days and mate, they lay eggs in squares that have = reached=20 at least the "one-third grown stage" (approximately 1/4 inch in = diameter). Egg=20 laying may occur in smaller squares; however, sufficient feeding = material is not=20 available for a high percentage of larvae to develop to the adult stage. = Late in=20 the season eggs may be laid in small bolls, but squares are preferred.=20

3D"Boll It takes the eggs 2.5 to 5 days to hatch into the grublike = larva=20 that feeds inside the square or small boll. After larval development = begins the=20 infested square turns yellow, bracts open or flare and the fruiting form = falls=20 off the plant. The larva feeds for 7 to 14 days before pupating inside = the=20 square or small boll. During the next 4 to 6 days the pupal stage = changes into=20 an adult boll weevil. The newly developed adult eats its way out of the = square=20 or small boll and feeds on other fruiting forms for about 5 days. During = this=20 time the weevil mates and females begin to lay eggs. The entire cycle = takes 16=20 to 18 days under ideal conditions. Six or seven generations may be = produced each=20 year with each female having the capability of laying approximately 200 = eggs.=20

Evidence of infestation. Although adult boll weevil feeding = causes=20 little damage, it indicates the presence of weevils and that egg laying = will=20 soon follow. There are distinct differences between feeding and = egg-laying=20 punctures.=20

As weevils feed, a small cavity is formed at the site of feeding. If = a female=20 weevil determines that the feeding site is suitable for egg laying, she = enlarges=20 the cavity slightly and inserts her ovipositor (egg-laying tube) to = deposit a=20 single egg in the cavity. When the ovipositor is withdrawn she secretes = a sticky=20 substance that covers the cavity. This sticky secretion hardens to form = a=20 wart-like protuberance that can be easily seen and felt. Feeding = punctures=20 usually have no sticky covering and therefore no wart-like protuberance. =

Nature of damage. Most of the damage is due to larval = development=20 inside fruiting forms. Feeding larvae eventually cause cotton squares = and small=20 bolls to shed or damage developing lint in larger bolls. Heavily = infested cotton=20 may produce much foliage but few mature bolls.

Cotton fleahopper (Pseudatomoscelis seriatus = [Reuter])=20

Identification. The adult cotton fleahopper is approximately = 1/8=20 inch long. It is flat with an elongated, oval outline and prominent = antennae.=20 The body usually is yellowish-green, although it may be white or yellow = with=20 minute black hairs and spots on the upper surface. After feeding, the = immature=20 stage is pale green with prominent, often reddish eyes. Other parts of = the body=20 also may be reddish. Cotton fleahopper eggs are yellowish-white, about = 1/30 of=20 an inch long and are inserted under the bark of small stems.

Biology. The cotton fleahopper overwinters in the egg stage, = primarily in wild hosts such as woolly croton (Croton capitatus), = cutleaf=20 evening primrose (Oenothera laciniata), showy sundrops (Oenothera = speciosa),=20 wooly tidestromia (Tidestromia lanaginosa), spotted [horsemint] beebalm = (Monarda=20 punctata), lemon [horsemint] beebalm (Monarda citriodora) and silverleaf = nightshade (Solanum elaeagnifolium). At 80=B0 F., eggs hatch in about 11 = days, and=20 the young nymphs feed on tender vegetation. They usually molt five times = and=20 require 14 to 15 days to mature into the reproductive adult cotton = fleahopper.=20 Usually there are six to eight generations per year. Of these, only one = to three=20 occur in the cotton field. Early in the spring, fleahoppers build up = large=20 numbers on alternate weed hosts. As these hosts mature and become less=20 succulent, the cotton fleahopper searches for more preferred hosts. If = cotton is=20 present, fleahoppers colonize in it and feed on leaf and fruit buds. = Some=20 fleahoppers stay in cotton fields as long as cotton plants are lush and=20 succulent.=20

3DCotton=20 Nature of damage and evidence of = infestation. The cotton fleahopper feeds on anthers of small = squares and=20 sucks sap from leaf buds. This feeding causes squares to die and turn = brown,=20 resulting in a "blasted" appearance. When fleahoppers are abundant, = heavy fruit=20 loss may occur on preflowering plants. The cotton fleahopper prefers = terminal=20 bud clusters including young leaves and tiny squares. The piercing, = sucking=20 habit of nymphs and adults interferes with normal growth patterns in = cotton.=20 Feeding punctures stimulate the plant to produce shorter main stem = internodes,=20 more nodes and spindly branches or "suckers" from the lower parts of the = plant.=20 The fleahopper injects saliva when feeding, but its effects are only = local.=20 There is no evidence that fleahoppers transmit plant diseases or toxic=20 substances other than digestive enzymes.

Bollworm (Heliothis zea [Boddie]) and Tobacco budworm = (Heliothis=20 virescens [Fabricius])

Identification. Bollworms and tobacco budworms pass through = four=20 developmental stages--egg, larva, pupa and adult. The adult bollworm is = a robust=20 moth with a wing span and body length of approximately 1 =BD inches. = Fore wings=20 vary from light brown or tan to reddish-brown and are 3D"Adult marked with dark areas near the tip and a dark spot usually = near the=20 center. The hind wing of the bollworm moth is white to light tan with an = irregular dark band on the outer hind margin. The adult tobacco budworm = is=20 similar but with a slightly smaller wing span and body length of about 1 = 1/4=20 inches. The fore wing of the tobacco budworm usually is light olive = green with=20 three or four light-colored, oblique bands. In the larval stages, = bollworms and=20 tobacco budworms look alike. Their size varies from very small (1/16 = inch) when=20 hatched up to 2 inches long as the larva reaches maximum size.=20

3D"Adult Tobacco budworm larvae have a tooth-like projection on = the inside=20 surface of the mandibles and fine short hairs on the first, second and = eighth=20 abdominal projection (tubercle) which bear a single, prominent spine. If = the=20 projection and hairs are absent, this indicates a bollworm. A microscope = or hand=20 lens is necessary to observe these characteristics.=20

3D"Bollworm The larval color of both species varies from a uniform = light green to=20 shades of green or brown, usually with stripes running the length of the = body.=20 It is impossible to distinguish between the two species by coloration. = Bollworm=20 and tobacco budworm eggs are similar. When recently laid, they are = whitish=20 colored and approximately the size of a pin head. Eggs are hemispherical = in=20 shape, resembling an inverted cup, with ridges running along the side = from the=20 top center to the point of attachment on the plant. They are usually = deposited=20 singly about the plant, and as they mature, usually within 2 to 5 days, = they=20 become tannish-brown. The darkening indicates the approaching hatch.=20

3D"Bollworm =20

Biology. The bollworm and tobacco budworm overwinter in the = pupal=20 stage in the soil near host plants. They pupate in the upper 5 inches of = the=20 soil. With adult emergence in the early spring, they commonly complete = several=20 generations on hosts other than cotton. These alternate hosts include = legumes,=20 corn and sorghum (bollworm only) or noncrop, native vegetation.=20

Moths enter cotton fields and feed on plant exudates (usually = nectar). Eggs=20 require 2 to 5 days to hatch. Larvae feed on fruiting forms for 14 to 18 = days.=20 After feeding ceases, larvae burrow into soil around plants and pupate = where=20 they spend 12 to 18 days. Upon emergence as adults, moths spend 2 to 5 = days=20 feeding on nectar from nectaries before egg laying begins again. The = complete=20 life cycle may take 35 to 50 days depending on weather conditions.=20

Nature of damage and evidence of infestation. The first = indication=20 of bollworm or tobacco budworm infestation may be newly deposited eggs = in the=20 terminal area of cotton plants. Eggs usually are laid on the upper = surface of=20 tender foliage and other tender parts including bracts, blooms, small = squares=20 and bolls. Newly hatched larvae usually begin feeding on tender leaf = surfaces=20 and other tender vegetation before attacking fruiting forms. Smaller = larvae feed=20 on smaller fruiting forms and generally move progressively to larger = fruiting=20 forms as they grow.

Larvae tunnel into small squares and terminal buds leaving holes that = range=20 in size from very small up to the diameter of a pencil. The entry site = usually=20 is at the base of fruit. In most cases, frass or excrement is seen at = the=20 feeding site. This frass is distinguishable from that of the boll weevil = because=20 it is slightly larger and is dark brown; boll weevil frass contains silk = strands=20 and is yellow or orange.
3D"Boll =20

Feeding damages or destroys the squares, blooms and bolls. Injured = squares=20 flare and drop from plants usually within 5 to 7 days. Large larvae feed = on=20 bolls, squares and pollen in open flowers. They may even devour the = contents of=20 large bolls. Worm-damaged bolls frequently are lost to boll rot even if = not=20 eaten completely. Larvae may "top" young plants by devouring the = terminal. This=20 often delays plant growth and may cause abnormal, nonproductive growth.=20

OCCASIONAL PESTS

Occasional pests are those whose populations increase to damaging = levels only=20 infrequently. In general they do not cause economic loss on a yearly = basis.=20 Occasional pest flare ups are often a result of disruptions in natural = control.=20

Beet armyworm (Spodoptera exigua [Hubner])

Identification. The adult stage of the beet armyworm has a = wing span=20 of 1 to 1 1/4 inches, and the fore wing is grayish-brown with two yellow = spots=20 near the center. The hind wing is a translucent white with narrow brown = borders.=20 The immature stage of the beet armyworm is a distinctive green larva up = to 1 1/4=20 inch long. There is a conspicuous black dot on each side of the second = body=20 segment behind the head. The female lays masses of about 80 eggs = beginning early=20 in the spring. These eggs are covered with hairs and scales from her = body.=20

Biology. The female lays an average of 500 to 600 eggs over = a 4- to=20 10-day period. These eggs take 2 to 5 days to hatch. The larvae then = feed for=20 about 3 weeks and pass through five instars (growth stages). As small = larvae=20 they may spin a light web over the foliage. Small larvae feed in groups = for=20 several days but later spread out and become solitary feeders. After = larval=20 feeding, 3D"Beet pupation takes place in the upper 1/4 inch of the soil in = a cell=20 formed by gluing soil particles and trash together with a sticky = secretion. This=20 entire life cycle from the egg to adult requires 30 to 40 days, = depending on=20 weather conditions. Beet armyworms overwinter in the pupal stage.=20

Nature of damage and evidence of infestation. The most = obvious=20 indication of infestation is the characteristic egg masses coupled with=20 defoliation by the larval feeding. Beet armyworms also feed on fruiting = forms of=20 cotton and occasionally may cause severe damage. Larvae often feed on = the bracts=20 causing little or no damage.

Brown cotton leafworm (Acontia dacia Drucel)

Identification. The adult stage of the brown cotton leafworm = is=20 sexually dimorphic, meaning males and females do not resemble one = another. The=20 fore wing of the female is gray at the base, bordered by an irregular = dark band=20 terminated with a white line. Each wing has a conspicuous reddish-brown = spot=20 near the trailing edge. The fore wing of the male is pale gray in the = basal area=20 and is bordered by an evenly curved white line extending obliquely = across the=20 wing. Here it joins a similar transverse line that separates the black = tip from=20 the white area. The average wing span of both male and female is about 1 = inch.=20 This is slightly smaller than the cotton leafworm.=20

The larval stage of the brown cotton leafworm is reddish-brown = mottled with=20 white. The larva is smooth and has a protuberance on the top of the = eighth=20 abdominal segment. Eggs are shaped like a football, bluish-green and = about the=20 size of a bollworm egg. They can be distinguished from the bollworm egg = by the=20 semitransparent ribs of varying lengths and the distinctive transparent = ridges,=20 resembling a band or belt. This insect pupates on the surface or in the = top 1/4=20 inch of the soil. The cocoon is capsule-like and made up of soil = particles,=20 plant material and silken fibers spun by the larva. 3D"Brown =20

Biology. Eggs, laid in the spring, hatch in about 3 days. = Larvae go=20 through several growth stages during a 12-day span. The pupal stage = lasts about=20 15 days. Therefore, the life cycle requires about 30 to 35 days. Adult = moths lay=20 eggs 3 to 4 days after emerging from the pupal case.=20

Nature of damage and evidence of infestation. The brown = cotton=20 leafworm is a foliage feeder. Damage in the early stage of an = infestation is=20 characterized by a buckshot pattern of holes in the leaves. Later, and = under=20 more severe attack, the plants may be defoliated completely.

Cabbage looper (Trichoplusia ni [Hubner]) 3D"Cabbage =20

Identification. The adult cabbage looper is a grayish brown, = mottled=20 moth approximately I inch long with a wing span of 1 =BD inches. There = is a=20 characteristic silvery spot resembling a "V" or a figure "8" with an = open end at=20 the top near the center of the fore wing. The hind wings are pale brown = but=20 darker toward the edge of the wing. The larva, when fully developed, is = about 2=20 inches long and light green with several white lines extending the = length of the=20 body. The larvae have three pairs of slender legs near the head and = three=20 thicker, club-shaped prolegs behind the middle toward the rear. The = middle half=20 of the body is without legs. Usually there are two light

3D"Cabbage 3D"Foliage
stripes on the top and one = stripe on the=20 side of the body. The body of cabbage loopers tapers to a small head. = Cabbage=20 looper eggs are light green and slightly flat. They lack the distinctive = ridges=20 of bollworm and tobacco budworm eggs. The eggs are deposited singly on = the=20 undersurface of leaves.=20

Biology. The cabbage looper overwinters in a cocoon and = usually=20 remains attached to the plant material upon which it has fed. Upon = emergence in=20 the early spring, adults feed for a short time and begin laying eggs. = One adult=20 moth may lay 200 to 350 eggs. The egg stage lasts 2 to 3 days. Larvae = feed for 2=20 to 4 weeks before changing into pupae in silk cocoons attached to = plants. The=20 pupal stage lasts 12 to 14 days before adult moths emerge. The complete=20 generation requires about 35 days.=20

3D"Cabbage Nature of damage and evidence of infestation. = Cabbage=20 loopers are principally foliage feeders. An infestation is characterized = by a=20 buckshot or tattered appearance in the leaves. Larvae have a = characteristic=20 movement from which the name is derived. Sometimes referred to as = "measuring,"=20 the larvae move about the plant by reaching out with the front part of = their=20 body, humping up the middle and pulling the rear part of their body = forward.=20 They may eat the area between leaf veins, occasionally creating a = net-like=20 appearance. Infestations may be greater along field margins. Cabbage = loopers are=20 seldom an economic pest; however, severe defoliation of cotton plants = may cause=20 problems with boll maturation. Severe defoliation before bolls mature = reduces=20 yields drastically. Any partial defoliation occurring later in the = season has=20 little effect on yield.=20

Cotton aphid (Aphis gossypii = Glover)=20

3D"Cotton

3D"Sooty
Identification. = Both the adult=20 cotton aphid and the immature stages are soft-bodied, sucking insects. = They=20 range from light yellow to dark green and in many cases are almost = black. Early=20 in the season they are a darker color when feeding on new growth of = cotton=20 terminals. Later in the season, when their feeding is restricted to the=20 underside of mature leaves, they are a lighter, yellowish color and are = smaller.=20 The immature or nymphal stage looks like the adult stage, only smaller. = Most=20 adults are wingless. There are no eggs as the young are born alive. This = aphid=20 should not be confused with the cowpea aphid (Aphis craccivora Koch). = The adult=20 of the cowpea aphid is shiny black with white patches on the legs. The = nymphs=20 are ash-gray.=20

Biology. The biology of the cotton aphid is unique. All = adults are=20 females. These females live and produce young on growing plants year = round.=20 Aphids reproduce rapidly. One female may produce as many as 80 young = females=20 that mature within 8 to 10 days. Thus, it is possible for aphids to have = as many=20 as 50 generations per year. These generations also occur as frequently = as every=20 5 to 7 days under optimum conditions.=20

3D"Cotton Nature of damage and evidence of infestation. = Aphids are=20 easily observed on cotton plants. They commonly appear on the underside = of=20 leaves where they suck the sap, causing leaves to curl and sometimes = shed.=20 Cotton aphids cause leaves to curl downward and "cup under, " which is = in=20 contrast to the upward curling caused by thrips. Aphids excrete = honeydew, a=20 sticky substance easily seen on cotton plants. A black sooty mold may = grow on=20 the honeydew during periods of high humidity. Feeding aphids may cause = malformed=20 plants or even kill seedling plants. If aphid populations are high at = harvest,=20 lint may become sticky with honeydew and interfere with ginning and = spinning.=20 Aphids serve as a very important food source for natural enemies of = other cotton=20 pests.=20

Cotton leafworm (Alabama = argillacea=20 [Hubner])

3DCotton=20 Identification. The adult cotton = leafworm=20 is a tan to brownish moth with a few darker, wavy, transverse bars on = the fore=20 wings. There may be an olive gray or purple tinge to the coloration. The = distinctive characteristics of the cotton leafworm adults are the = undulating=20 reddish lines across the front wings and the oval dark spot near the = center of=20 each fore wing. The wing span is about 1 =BD inches. Cotton leafworm = larvae are=20 smooth, light to dark yellowish-green and marked with three narrow white = stripes=20 down the back and one along each side.=20

The larvae have four black dots that form a square on the top of each = body=20 segment, similar to a double four domino. Each black spot has a black = spine, and=20 the center is surrounded with a white ring. This gives it the domino=20 characteristic. The size varies from very small to 1 =BD inch in length. = Larvae=20 are pale when small and change to a brighter color as they mature. Eggs = of the=20 cotton leafworm usually are seen on the underside of leaves. They are=20 bluish-green when first laid, flat and barely as large as the head of a = pin.=20

Biology. The cotton leafworm is a tropical species and does = not=20 overwinter in the U.S. In the spring, adult moths fly in from the = tropics or are=20 blown in on prevailing winds. They begin laying their eggs on cotton = leaves.=20 These eggs are scattered on the underside of the leaves and laid singly. = Eggs=20 hatch within 3 days and the young larvae begin feeding. Larvae pass = through five=20 to six growth stages per generation, each a different length. During = optimum=20 conditions and in mid-summer the larval stage lasts approximately 15 = days. The=20 larvae of cotton leafworms web up with a boll or roll of leaves forming = flimsy=20 cocoons for protection just before pupation. Pupation, which occurs on = the=20 cotton plant, takes approximately 6 to 10 days. Newly emerged adults lay = eggs=20 within 2 to 4 days.=20

3D"Cotton Nature of damage and evidence of infestation. = Feeding=20 patterns of cotton leafworms are characteristic because they feed on the = underside of leaves between the veins, skeletonizing them. The = skeletonizing of=20 leaves causes a reduction in photosynthetic potential. Although this = insect is=20 essentially a leaf feeder, it can attack squares and small bolls. The = cotton=20 leafworm in Texas is a late-season pest that often aids defoliation just = before=20 harvest.

Cotton leafperforator=20 (Bucculatrix thurberiella Busck)=20

Identification. The adult cotton leafperforator is a small, = slender,=20 usually white moth with black markings on the fore wings. The wing span = and body=20 length are approximately 2/5 to =BD inch. The larva is a dull amber = green with two=20 rows of black spots and distinct white projections (tubercles) on its = back. The=20 larva reaches a maximum length of =BD inch. The pupa is a cocoon of = silk-like=20 material on the stalks of cotton plants. These pupal cocoons are white,=20 approximately =BD inch in length and arranged longitudinally on the = plant. Eggs=20 usually are laid singly on cotton leaves.=20

Biology and evidence of infestation. After eggs hatch, early = instar=20 larvae make a winding, randomly patterned mine in leaves. The larvae = feed in the=20 mines through the third instar. At the fourth instar they emerge from = the mine=20 as free feeders. Damage from these free-feeding larvae skeletonizes the = leaf.=20 Cotton leafperforators have a characteristic horseshoe-shaped stage that = marks=20 the time between the fourth and fifth instar larva. This resting stage = is in the=20 form of a small, grayish horseshoe and is obvious on the underside of = the leaf.=20 When cotton leafperforator larvae emerge from the horseshoe stage they = continue=20 to feed freely. Mature larvae spin a fine web, drop to the ground and = move along=20 the ground to the base of cotton or other plants in the area. There the=20 characteristic slender, ribbed, white cocoon is constructed. The cotton=20 leafperforator is most frequently a problem in hot, dry climates.

Nature of damage. The main damage done by the cotton = leafperforator=20 involves interference with normal boll maturation. Severe infestations = halt=20 normal growth and respiration of the cotton plant and effectively stop = the=20 maturing process. Heavy populations are commonly associated with = multiple=20 insecticide applications that destroy parasites and predators. = Fortunately,=20 infestations usually occur late in the season when the crop is mature.=20

Cotton square borer (Strymon = melinus=20 [Hubner])

Identification. The adult cotton square borer is a slate or=20 bluish-gray butterfly. It has two large black spots on the wings with = smaller=20 reddish-orange spots above these at the back edge of the hind wing. = There are=20 usually two or three thin tails on the hind wing. The underside of the = wings is=20 lighter gray with a white streak along the edge. The wing spread of the = cotton=20 square borer adult is approximately 1 inch. The larval stage is a = velvety, light=20 green, slug-shaped worm. The body is covered by a succession of oblique = lines=20 running the width of the body segments. Cotton square borer eggs are = pale green=20 and laid singly on plants. 3D"Cotton =20

Biology. The adult begins laying eggs early in the spring. = The eggs=20 take approximately 6 days to hatch; then the small larvae feed on cotton = squares=20 for approximately 20 days. After the feeding period the insect spends = about 10=20 days in the pupal stage.=20

Nature of damage and evidence of infestation. Presence of = adult=20 butterflies usually is the first indication of cotton square borer = activity.=20 However, hollowed out squares with almost perfectly round entrance/exit = holes=20 also may be observed. Unlike bollworm or tobacco budworm damage, there = is no=20 frass present with cotton square borer damage.=20

Cotton stainer (Dysdercus = suterellus=20 [Herrick-Schaffer])

Identification. The adult cotton stainer is a "true bug" = with=20 piercing, sucking mouthparts. The head and pronotum are bright red; the=20 remainder of the body is dark brown crossed with pale yellow lines. The = length=20 is =BD inch or slightly longer. Immature stages are smaller but resemble = adults=20 without wings. Cotton stainer eggs are small, pale and laid singly on = cotton=20 plants or dropped on the ground near cotton plants.=20

Biology. Eggs hatch in 5 to 8 days. There are five instars = in the=20 immature stage which lasts about 21 to 35 days. The first instar = immatures are=20 usually found congregating near the egg shell after emergence. The = second and=20 third instars feed gregariously on the bolls. Later instars wander = freely over=20 the plant.=20

Nature of damage. The cotton stainer is mainly a pest of = long staple=20 cotton. It damages developing bolls by puncturing seeds and causing = plant sap to=20 exude from the feeding site. The plant sap stains the lint an indelible = yellow=20 color. Feeding by the cotton stainer also interferes with the bolls' = natural=20 development.=20

Cutworms (Agrotis spp.)

3D"Cutworm
Identification. The = adult cutworm=20 is a brown to gray robust moth. The wing span varies from I to 1 =BD = inches.=20 Several species infest cotton. Cutworm larvae characteristically curl up = into a=20 C-shaped posture and regurgitate readily when disturbed. The larvae are = often=20 shiny or glossy and always have four pairs of prolegs.=20

Biology. Most cutworms overwinter in the larval stage, but = some=20 overwinter as pupae or adults. The eggs are laid on grass or soil in low = spots=20 of the field. They normally hatch in 2 to 5 days. 3D"Cutworms The average time of larval feeding is 2 to 3 = weeks. The=20 pupal stage remains in the soil for 7 to 8 days. About 35 days normally = are=20 required to complete a generation.=20

Nature of damage and evidence of infestation. Cutworms = usually cut=20 off plant stems at the soil surface. Stand reduction may be more visible = in=20 field margins and low lying weedy areas. Some cutworm species climb up = plants=20 and feed on leaves. Cutworm damage can be severe enough to require = replanting on=20 rare occasions.

Fall armyworm (Spodoptera = frugiperda=20 [J.E. Smith])=20

3D"Fall Identification. The adult fall armyworm is a = dark=20 brownish-gray, mottled moth with oblique markings near the center of the = fore=20 wing. There is an irregular white or gray patch near the wing tip. = Female moths=20 are darker than males. The hindwing is white in females with a pearly or = pink=20 luster and a brown border. The wing span of the fall armyworm is = approximately 1=20 =BD inches. The larva of the fall armyworm, when newly hatched, is white = with a=20 black head. As feeding progresses, the larva becomes darker. Young fall=20 armyworms often curl up on a leaf or suspend from a silken thread. As = the larvae=20 mature they turn greenish-brown with a white line below the top of the = back,=20 usually a brownish-black stripe above the midline and a pale stripe with = a=20 reddish-brown tinge below. Mature larvae are about 1 =BD inches long. = There is a=20 prominent white inverted "Y" on the front of the head. Fall armyworm = larvae can=20 be distinguished from beet armyworms by the presence of black hairs on = the body.=20 Fall armyworm eggs are similar to those of the beet armyworm. They are = laid in=20 masses of 50 to 100 and covered with hairs and scales from the female's = body.=20

Biology. The fall armyworm is another tropical insect which = is=20 apparently unable to overwinter in the United States except during mild = winters.=20 In the early spring swarms of moths are blown up or fly northward from = tropical=20 areas. Approximately 3 to 5 days are required for the eggs to hatch.=20 Developmental time of the larval stage takes 2 weeks to a month. = Pupation occurs=20 in the ground near the cotton plant and usually requires 1 to 2 weeks. = The adult=20 begins laying eggs after 3 or 4 days and lays about 150 eggs a day for 8 = to 10=20 days.=20

3D"Fall Nature of damage and evidence of infestation. = When=20 abundant, they generally eat all available foliage and then crawl in = armies to=20 adjoining fields. Fall armyworms are general feeders and do not confine=20 themselves to cotton.=20

Fall armyworm populations can build up large numbers in cotton = fields, and=20 defoliation may be quite severe on rare occasions. The greatest damage = comes=20 from the topping of plants. Branches may be cut off and sometimes the = stalks may=20 almost be completely severed a foot from the top. They also tend to feed = on=20 squares, blooms and bolls on the flower part of the plant. Several worms = may be=20 found on individual plants. They are an occasional cotton pest where = grasses=20 grow in or near cotton fields.=20

Grasshoppers -- Lubber grasshopper = (Brachystola magna [Girard]); Differential grasshopper (Melanoplus spp.) =

Identification. Adult grasshoppers vary in appearance, but = all have=20 well developed back legs for jumping. The immature stage is similar to = the adult=20 but does not have wings. Grasshopper eggs are laid in the soil in a = cemented=20 mass or pod. Depending on the species, there may be 8 to 25 eggs in each = pod.=20 These pods usually are distinctive and are used in species = identification.=20

Biology. The biology of the different grasshoppers varies=20 considerably. However, in most species the female lays eggs in the soil = where=20 overwintering takes place. Some species overwinter as 3D"Grasshopper partially grown nymphs or as adults. Upon = hatching in the=20 early spring, grasshoppers usually require 40 to 60 days of feeding = before=20 reaching reproductive maturity. Seldom is there more than one generation = per=20 year.=20

Nature of damage. Grasshoppers may become problems in cotton = fields=20 when wild or cultivated host plants are depleted near fields. When = infestations=20 are heavy, grasshoppers cause severe defoliation by feeding on foliage = and=20 tender plant parts. Their damage usually is confined to areas along = field=20 margins where alternate host plants are near.

Tarnished plant bug (Lygus lineolaris = P. de=20 Beav.)=20

3D"Tarnished

Identification. Adult tarnished plant bugs are about 1/4 = inch long=20 and approximately half that wide. They are flat and have an oval = outline. They=20 are similar to cotton fleahoppers in shape; however, they usually are = about two=20 to three times larger than the cotton fleahopper. They are generally = brown in=20 color with splotches of white, yellow and reddish-brown. Along the side = of the=20 body on the hind part of the wing there is a clear, black-tipped = triangle. This=20 description varies from the lygus bug, typically found in West Texas, in = that=20 its color ranges from pale green to yellowish brown. There is a = conspicuous=20 light-colored triangle on the back of the Lygus bug (Lygus hesperus = Knight)=20 between the wings. When first hatched, tarnished plant bug nymphs are = pale green=20 and have an orange spot in the middle of the abdomen. After feeding = begins,=20 nymphs become darker green and black spots become evident on the body = segment=20 (thorax) between the head and abdomen. The whitish egg is slightly = curved,=20 elongated and has a lid on its tip.=20

Biology. In most areas tarnished plant bugs overwinter as = adults.=20 Mature adults lay eggs in the cotton plants; these hatch in about 8 = days. The=20 immature nymphal stage develops in approximately 11 days. Each = generation is=20 completed in 20 to 30 days in the summer, and there may be more than one = generation per year.=20

Nature of damage and evidence of infestation. Like cotton=20 fleahoppers, most tarnished plant bugs occur on other host plants in = areas=20 surrounding cotton fields. They feed by inserting their 3D"Tarnished mouthparts into the tender plant parts and = sucking sap.=20 Feeding on squares and small bolls usually causes fruit shed. The most = important=20 period of plant susceptibility is during the first 3 weeks of squaring = with the=20 first week being the most critical. During heavy infestations fruit = removal can=20 be excessive and of economic importance. Occasionally, damaged squares = and bolls=20 do not shed. Although squares may bloom, floral parts are damaged and = show=20 evidence of feeding. If an injured boll does not shed, it opens = abnormally and=20 may have lint damage.

Omnivorous leafroller (Platynota = stultana=20 Walsingham)=20

Identification. The adult stage of the omnivorous leafroller = is a=20 small moth approximately =BD to 3/5 inch long. The fore wing has a dark = brown=20 inner half and a yellowish-brown outer half. Mouthparts that extend in = front of=20 the head give the appearance of a beak. The immature stage is a = yellowish-white=20 caterpillar with a brown head and shield behind the head. As the larva = matures=20 the coloring changes to yellowish-green or brownish-green with a broad=20 transparent stripe down the top of the back. The larva is approximately = 3/5 inch=20 long. The green eggs are laid in flat clusters overlapping one another = like fish=20 scales. The egg mass is cemented to the plant with a secretion from the = female's=20 reproductive glands.
3D"Omnivorous =20

Biology and evidence of infestation. The adult female = omnivorous=20 leafroller may lay more than 300 eggs during her life span. These eggs = take only=20 a few days to hatch. The dark-headed larvae begin making silken ladders = on which=20 they move about and feed. The larva has a peculiar habit of concealing = itself by=20 rolling a leaf around its body. Some larvae develop completely in a = single leaf=20 roll, while others may build several. After about 25 days of feeding = pupation=20 takes place within the leaf roll. The adult emerges about 5 days later.=20

Pink bollworm (Pectinophora = gossypiella=20 [Saunders])

Identification. The adult pink bollworm is a =BD inch = grayish-brown=20 moth with a 2/5 inch wing span. When it rests, the moth's wings are = folded close=20 to the body. These wings are covered with poorly defined black spots and = have=20 blackish tips with a broad band near the tip. Wing margins are fringed = with=20 hairs. The immature pink bollworm is a small larva with a yellowish = brown head.=20 It may be as long as =BD inch just before pupation. Young larvae are = white and=20 turn pink with age. The pink coloration occurs in dark transverse bands = running=20 across or covering the body. Pink bollworm eggs are greenish-white and = are laid=20 on the stems and squares and occasionally on terminal buds. They are = most often=20 found between the calyx and boll wall. Eggs are usually deposited = singly.=20

Biology. Eggs normally hatch within 4 to 5 days. Developing = larvae=20 bore into and feed on the developing flower or into a boll to feed on = the seed.=20 Larvae feed for 10 to 14 days. They usually leave the fruit and pupate = in the=20 soil, taking about 8 days to transform into an adult. In late summer and = fall=20 many larvae do not pupate immediately upon completion of the feeding = period but=20 remain inside the boll. These are the overwintering stages that give = rise to the=20 first generation the following season. Overwintering usually takes place = in seed=20 inside bolls or inside two hollowed out seeds with a pupal case = connecting them.=20 These "double seeds" are found when cotton is ginned.=20

3D"Pink Nature of damage and evidence of = infestation.=20 Early in the season, infested squares continue growing and producing = blooms.=20 These infested blooms may have the petals tied together with silken = threads,=20 commonly termed "rosetted." The rosetted flowers do not open normally. = After=20 bolls are present they become the preferred egg-laying site. The = developing=20 larvae tunnel through the boll wall until they reach the lint. They cut = through=20 the lint fiber as they move from seed to seed. The burrowing activity = stains=20 lint; destroys fibers; and reduces seed weight, vitality and oil = content. Pink=20 bollworms cut holes in boll walls as they leave bolls for pupation. = These holes=20 may become infected with boll-rotting organisms. During severe = infestation, many=20 bolls that might otherwise have been harvested are rendered unpickable.=20

Saltmarsh caterpillar (Estigmene = acrea [Drury])=20

3D"Saltmarsh
Identification. The = adult saltmarsh=20 caterpillar is a large, light-colored moth. The abdomen usually is = orange to=20 yellow and may have black spots on the top. Male and female moths differ = in the=20 way their wings are colored. In the male the fore wing is white and the = hind=20 wing is orange to yellowish. In the female both wings are snowy white on = the top=20 and yellow below. The wing surfaces of both sexes are marked with = variable black=20 spots. The wing span usually is 1 =BD to 2 inches. The larval stage of = the=20 saltmarsh caterpillar is sometimes called a "wooly bear," because the = body is=20 covered with long hairs giving it a wooly appearance. These larvae are = gray to=20 black to reddish brown with a dark head. The full sized larva reaches a = length=20 of 2 1/4 inches.=20

3D"Saltmarsh Biology. The saltmarsh caterpillar = overwinters=20 under plant debris in the pupal stage inside a fragile, silken cocoon = heavily=20 covered with interwoven hairs from the caterpillar's body. Upon = emergence in the=20 early spring, the female lays as many as 1,000 spherically shaped eggs = in=20 patches on the underside of host plant leaves. Newly hatched saltmarsh=20 caterpillar larvae are gregarious. They feed in large numbers on the = bottom side=20 of leaves. As the larvae mature they disperse and feed singly. The = larval stage=20 requires 30 to 48 days for completion; pupation may take up to 14 days. = There=20 usually are two to five generations per year.

Nature of damage and evidence of infestation. This insect = usually is=20 a problem on the edges of fields and in areas where alternate host = plants are=20 available. These pests can be severe defoliators when present in large = numbers.=20 Saltmarsh caterpillars may migrate in mas from one field to another like = armyworms.=20

Spider mites (Tetranychus = spp.)=20

3D"Spider Identification. Spider mites are not = insects.=20 They have eight legs in the adult stage instead of the six which is=20 characteristic of insects. The adults are so small they can barely be = seen with=20 the naked eye. They are usually red, but may be green, orange or straw = colored.=20 Casually observed, spider mites often look like dust particles on the = underside=20 of leaves.=20

Biology. Although spider mites attacking cotton are really a = complex=20 of several genera and species, they all have similar life histories and = biology.=20 Spider mites pass through egg, larval, nymphal and adult stages. They = overwinter=20 in the adult female stage in protected vegetation near fields. The = female lays=20 her eggs on the underside of leaves in early spring. Eggs hatch in 1 to = 3 days.=20 Freshly hatched larvae have only six legs and are transparent. They feed = for 2=20 to 3 days before molting (shedding their skin) and developing into = eight-legged=20 nymphal mites. Nymphs feed for 5 to 6 days before molting into = eight-legged=20 adults. Adults feed for 1 to 3 days before laying eggs. The entire life = cycle=20 requires only 6 or 7 days under optimum conditions and 11 days under = normal=20 field conditions. Spider mites have an unusual life history in that = fertilized=20 eggs produce females and unfertilized eggs hatch into males.=20

3D"Spider Nature of damage and evidence of = infestation.=20 Spider mite infestations should be suspected when small, yellow spots = appear on=20 the upper surface of cotton leaves. This stippled discoloration is = typical of=20 early feeding stages. These yellow areas often result in leaf reddening = as mite=20 feeding continues. Large numbers of mites cause leaves to die and = complete=20 defoliation can occur. The lower surface of an infested leaf usually is = covered=20 with a fine web and cast skins. Live spider mites can be observed = crawling=20 beneath the web on the underside of the leaf surface. Spider mites cause = more=20 harm to cotton plants during hot, dry weather and usually attack cotton = in the=20 latter part of the season. Mites damage cotton by feeding on the sap of = leaves,=20 stems and squares. Since they suck sap and chlorophyll from the leaves, = this=20 reduces the photosynthetic area and interferes with normal maturation of = the=20 cotton plant. Spider mite infestations generally begin on the edge of = fields and=20 then spread through the field. They frequently are a problem following=20 insecticide applications which kill their natural enemies.

Stink bugs (Acrosternum hilare = [Say]) and=20 (Chlorochroa ligata [Say])=20

3D"Green Identification. Two species of stink = bugs are=20 typically found in Texas cotton and may cause problems. The green stink = bug=20 (Acrosternum hilare [Say]) and the conchuela stink bug (Chlorochroa = ligata=20 [Say]) are most often involved. The adult stages of these two stink bugs = are=20 similar. They are shield-shaped, flat and vary in size around =BD inch = in length.=20 The green stink bug is bright green, while the conchuela stink bug is = dark brown=20 to black with a red border and a red spot on the tip of the abdomen. The = immature stage of the stink bug is similar to the adult but lacks wings. = Stink=20 bug eggs have a distinctive barrel shape and usually are laid in = clusters on=20 stems and leaves. These egg masses resemble many barrels lined up in = rows.=20

Biology. The biology of these two species follows the same = pattern.=20 Eggs are laid on the host plant and require only a few days to hatch. = When=20 climatic conditions are favorable, a generation of these insects may be=20 completed in about 6 weeks.=20

3D"Stink Nature of damage and evidence of = infestation.=20 Stink bugs seldom develop in large numbers on cotton. They feed by = inserting=20 their long, piercing mouthparts and sucking out the plant juices thereby = injuring squares and bolls. This may cause small bolls to fall from the = plants.=20 Although larger bolls commonly remain on the plant, feeding injury = results in=20 hardened, dry locks. At each feeding site there is a hardened spot. Lint = beneath=20 the feeding spot may be stained or reduced in grade. Boll-rotting = organisms also=20 are associated with the feeding of these insects. In Texas stink bugs = rarely=20 occur in damaging numbers. They are most likely to be found in damaging = numbers=20 late in the season.

3D"Conchuela = 3D"Stinkbug

Thrips (Thrips spp.) and (Frankliniella=20 spp.)=20

3D"Thrips 3D"Severe

Identification. Thrips are minute, slender, agile insects. = They=20 rarely exceed 1/15 inch in length and usually are yellowish-brown, = black, tan or=20 sometimes orange. Their mouthparts are cone-shaped, and their wings are = narrow=20 and fringed with hairs.=20

3D"Thrips Biology. Thrips appear on growing plants throughout the = year in=20 most areas. Eggs, inserted into the plant tissue by the female's sharp=20 egg-laying tube (ovipositor), hatch in about 6 days. There are two = larval stages=20 that require about 6 days for completion.=20

The prepupal and pupal stages require an additional 4 days. Egg to = adult=20 development requires about 16 days. The average life span of a mated = female is=20 about 35 days. Fifty or more eggs may be produced. Thrips can reproduce = without=20 mating. Mated females produce both males and females while unmated = females=20 produce only males. Nature of damage and evidence of = infestation.=20 Thrips are early season pests of seedling cotton. They may be a problem = under=20 cool, wet conditions when plant growth is slowed. They may be especially = numerous in cotton grown near maturing small grains. Thrips damage = cotton leaves=20 and terminal buds. Their feeding ruptures cells which cause stunted = plants and=20 crinkled leaves that curl upward. During severe infestations terminal = buds may=20 be destroyed, causing excessive branching of the plants and delaying = plant=20 growth. Also, the cotyledonary leaves take on a silvery appearance, = sometimes=20 termed "bronzing.

Whiteflies (Bemesia tabaci = [Gennadius])=20

3D"Adult 3D"Whiteflies
Identification. The = adult whitefly=20 is about 1/16 inch long and snowy white. These moth-like insects are = very active=20 and fly readily when disturbed. The immature stage is a flat, scale-like = insect=20 usually about 1/30 inch long. The immatures usually are found on the = underside=20 of leaves. Whitefly eggs are only visible under magnification.=20

Biology. Whiteflies are active throughout the year on = alternate=20 hosts. Adults mate immediately after emerging from pupae. Eggs are laid = on fully=20 opened leaves in the terminal. These eggs require about 5 to 6 days to = hatch.=20 The nymphs, also called crawlers, move short distances, but after they = insert=20 their mouthparts into the leaves they become immobile. They feed for 5 = or 6 days during the first three growth stages. = Another 5 or 6=20 days are spent in a pseudo-pupal stage where no feeding occurs for the = last 2=20 days. After this stage the adult emerges ready to produce eggs. The = adult female=20 lays an average of seven eggs a day for 3 weeks. Peak whitefly numbers = usually=20 occur early in the season and during high humidity periods.=20

Evidence of infestation and nature of damage. These tiny = snow white=20 insects are easily observable flying about the cotton plant. They = excrete=20 honeydew much like aphids. Infested leaves lack vigor, wilt and may turn = yellow.=20

Yellow-striped = armyworm=20 (Spodoptera ornithogalli [Guenee])

Identification. The adult yellow-striped armyworm is a large = moth=20 with a wing spread of 1 =BD to 1 3/4 inches. The fore wing is = blackish-brown,=20 occasionally with a reddish tint, or may be yellowish-brown with oblique = yellow=20 markings near the center. There may be bluish-gray markings at the apex = and the=20 hind angle of the fore wing. The wing veins usually are lighter colored. = Hind=20 wings usually are irridescent white with a narrow brown border. The = larval stage=20 is a smooth caterpillar with scattered short hairs. As with other = armyworms=20 found in cotton, the inverted "Y" on the front of the head is prominent. = The=20 yellow-striped armyworm has a pair of black, triangular spots below the = top=20 surface on each segment except those between the head and the abdomen. = There are=20 yellowish-brown or occasionally orange stripes with white lines along = each side.=20 These yellow lines give the yellow-striped armyworm its name. Eggs are = laid in=20 masses on leaves and are covered with hairs and scales from the female's = body.=20

3D"Yellow-striped Biology. This species overwinters in the pupal = stage.=20 After emerging in early spring, the female feeds for a couple of days = before=20 laying eggs. Eggs hatch in about 6 days. The larval stage takes = approximately 20=20 days to complete. Pupation usually takes about 14 days during the = summer. An=20 entire life cycle from egg to adult requires 35 to 45 days.=20

Nature of damage and evidence of infestation. The = yellow-striped=20 armyworm is a day-feeding insect. The brightly colored larvae are easy = to=20 recognize. They prefer foliage as the feeding site. They also bore into = squares=20 and bolls. They are general feeders and infrequent cotton pests, = although when=20 alternate hosts are not available they can defoliate cotton and damage = the=20 fruiting forms.

PREDACEOUS INSECTS AND SPIDERS

W.L. Sterling=20

In Texas, native predaceous insects and spiders are the dominant = biological=20 control agents of the bollworm (Heliothis zea [Boddie]) and the tobacco = budworm=20 (H. virescens [Fabricius]). Native parasites are also valuable natural = enemies.=20 Other predators such as birds, lizards, snakes, toads, frogs, mice, = skunks,=20 armadillos, etc., also prey on these insect pests, but their = dependability and=20 efficiency generally are much lower than that of parasites, predaceous = insects=20 and spiders in cotton fields.=20

Native biological control organisms, along with other factors such as = heat,=20 cold, moisture, etc., often maintain cotton pests below numbers that = cause=20 economic crop losses. About 60 percent of the cotton in Texas is grown = without=20 requiring chemical insecticides, thus dependence is entirely on = biological and=20 physical control. Researchers are working to establish economic = thresholds=20 (action levels based on pest densities at which protective action is = needed to=20 prevent unacceptable losses) and to establish predator densities at = which no=20 action (inaction levels) is needed because the predators are sufficient = to=20 maintain pests below the economic threshold. It is essential to know = which=20 predators are most abundant and efficient and when they move to or from = the=20 field. A knowledge of predator effectiveness against individual pest = species=20 also is needed.=20

Predator densities and seasonal changes in populations presented in = the=20 following discussion are based on levels observed in East Texas cotton = fields=20 where broad spectrum insecticides are not used directly on the crop or = do not=20 drift into the field. The time of peak predator densities and average = densities=20 varies across Texas depending on prey availability, growth stage of the = cotton=20 plant, predator distributions and many other factors. The species and = number may=20 vary in some years and in some places.=20

The choice of predators for this discussion was based on their = relative=20 abundance in cotton and their importance as egg and small larval = predators of=20 bollworms or tobacco budworms. These predators often eat eggs and small = worms=20 before they develop into large worms that cause crop losses. In general, = small=20 predators feed on small prey such as eggs and small larvae, and larger = predators=20 kill large worms. Thus, identification of small, immature insect and = spider=20 predators, as well as adults, is important in pest management.=20

Most predators in cotton fields are general feeders that do not = depend on a=20 single pest species for food. Thus, if one prey species becomes scarce,=20 predators switch to another prey or may even resort to plant juices or = nectar to=20 survive. A complex of predator species can coexist and bring stability = to the=20 system. As any one pest species increases, bringing it out of balance = with the=20 system, predators switch to this new food source and again bring the = pest into=20 equilibrium.=20

Native predators are of great economic benefit to the cotton farmer. = They=20 voluntarily enter the cotton field, are self-multiplying and are = completely free=20 of cost.=20

More than 600 insect and spider predator species have been recorded = in cotton=20 fields. The 16 most important species are included in this publication.=20

Insects

The abundance and importance of insect predators vary considerably = from year=20 to year and from place to place. In any particular locality certain = insect=20 predators not mentioned here may be dominant. However, current evidence = suggests=20 the following eight insects play a dominant role in regulating bollworms = and=20 tobacco budworms in many cotton fields.=20

Minute pirate bugs = (Orius=20 spp.)=20

3D"Minute Two species of pirate bugs are found on cotton in Texas. = In eastern=20 Texas the insidious flower bug (Orius insidiosus [Say]) is = often the=20 dominant species. In West Texas and the Lower Rio Grande Valley it is = often=20 replaced by the minute pirate bug (Orius tristicolor [White]). = Both=20 species are black and white and measure less than 1/8 inch.=20

Pirate bugs usually are one of the most abundant cotton predators = averaging=20 three per meter (40 inches) of row with peaks of five to 10 per meter of = row in=20 mid-July in East Texas. They enter the cotton field very early in the = season,=20 reaching a spring peak in June or July and a fall peak in September. = They feed=20 on aphids, thrips, whiteflies, mites and bollworm-budworm eggs and small = larvae.=20 Reproduction occurs in cotton. The orange pirate bug nymphs also are = predaceous.=20

When bollworm or tobacco budworm eggs are plentiful, pirate bugs eat = about=20 one egg per day. They also kill and consume about one or two small worms = every 2=20 days.=20

Bigeyed bugs (Geocoris = spp.)=20

3D"Bigeyed Several species of bigeyed bugs are found in Texas = cotton; however,=20 the two most abundant species are G. punctipes [Say]), a large = grey=20 species, and G. uliginosus [Say]), a small black species. All = bigeyed=20 bugs found on cotton have distinctively broad heads, large conspicuous = eyes and=20 range in length from 1/8 to 1/4 inch.=20

In East Texas, bigeyed bugs peak in mid-July at two to five per meter = of row=20 while averaging about one per meter over the season. They enter cotton = fields as=20 soon as prey is available and remain throughout the growing season. Both = adults=20 and nymphs feed on mites, whiteflies, thrips, Lygus bugs and fleahoppers = while=20 consuming about two bollworm or tobacco budworm eggs per day under field = conditions. They also may feed on up to one small bollworm or tobacco = budworm=20 larva per day. They readily feed on various seeds and suck plant juices = but are=20 not considered harmful to the plant.=20

Reproduction takes place in the cotton field. Nymphs of G. = punctipes=20 are gray while G.uliginosus nymphs are solid black.=20

Ants (Solenopsis spp.)=20

3D"Red Many species of predaceous ants may be found on unsprayed = cotton in=20 Texas. Probably the most important are the fire ants (Solenopsis = species). In=20 areas inhabited by the red imported fire ant (Solenopsis = invicta=20 [Buren]), this ant specie often makes up 98 percent of all ants present. = In West=20 and South Texas where imported fire ants are not found, other species = such as=20 the tropical fire ant (S. geminata [Fabricius]) or in North = Texas the=20 southern fire ant (S. xyloni [McCook]) are important predators. = Worker=20 ants, generally found in cotton fields, are about 1/8 inch long, dark = red to=20 black, wingless and have a two-segmented waist (i.e., abdominal stalk).=20

Ants enter cotton when aphids start producing honeydew and may reach=20 densities of 10 to 20 ants per meter of row while averaging about eight. = Young=20 queens colonize cotton fields after rains. Ant numbers usually peak in = late June=20 and again in early August. Each ant may take about 0.3 bollworm or = tobacco=20 budworm eggs and 0.1 small worms per day. Since ants can hunt in packs, = they can=20 capture rather large prey and are excellent predators of immature boll = weevils.=20 Fire ants are relatively ineffective predators of most other predaceous = insects=20 and spiders.=20

Lady beetles = (Hippodamia=20 spp.)=20

3D"Convergent Of the many large lady beetles found in cotton, the most = numerous=20 and effective is the convergent lady beetle (Hippodamia = convergens=20 [Guerin-Meneville]). The adult generally is orange with 12 black spots = on the=20 wing covers (elytra). The orange and black larva also is predaceous. = Lady beetle=20 populations are related to aphid numbers, so lady beetles are most = abundant when=20 aphid populations are high.=20

Although large lady beetles are not as abundant as some of the other=20 predators, they apparently find and eat more bollworm-budworm eggs per=20 individual. Under field conditions they are capable of finding and = eating about=20 12 eggs per day. They also attack small-to-medium bollworm or budworm = larvae.=20

Cotton fleahoppers=20 (Pseudatomoscelis seriatus [Reuter])=20

3D"Cotton This insect is considered an important cotton pest = because its=20 feeding causes small squares to shed. However, the fleahopper also = should be=20 considered a predator on bollworm or tobacco budworm eggs. Under field=20 conditions the fleahopper may eat about one bollworm or tobacco budworm = egg per=20 day.=20

Fleahopper adults enter the cotton field when plants are very small. = As an=20 example in East Texas, their numbers peak in May or June and again in = September.=20 In some locations they may average about seven adults and nymphs per = meter with=20 peaks of 20 per meter. Since fleahoppers generally are considered pests = only on=20 preflowering cotton, they might be considered beneficial after = flowering. Both=20 adults and nymphs are predaceous but also serve as food for many other=20 predators.=20

Green lacewings = (Chrysopa=20 spp.)=20

3D"Green Although many species of green lacewings are present in = cotton=20 fields, the most abundant are the common green lacewing (Chrysopa=20 carnea [Stephens]) and the green lacewing (Chrysopa = rufilabris=20 [Burmeister]). Adults are about =BD to 3/4 inch long, and are = yellowish-green with=20 golden eyes and large, delicate, netted wings. The larvae grow to a = maximum=20 length of 3/8 inch, are alligator-shaped, have long sickle-shaped jaws = and are=20 mottled brown. Eggs are deposited on the end of long delicate stalks.=20

In East Texas green lacewings peak during mid-July at about one per = meter of=20 row, averaging one per 5 meters. Green lacewings often are more abundant = in=20 central and western parts of Texas. Movement begins in February or = March,=20 reaching a spring peak in May and a fall peak in September.=20

Common green lacewing adults are not predaceous, but the immatures = feed on=20 aphids, spider mites, whiteflies and bollworm or tobacco budworm larvae = and may=20 consume about one egg or small worm every 2 days.=20

Damsel bugs (Nabis = spp.)=20

3D"Damsel Several species of damsel bugs are found on Texas cotton. = One of=20 the most abundant species is the pale damsel bug (Nabis = capsiformis=20 [Germar]). Adult damsel bugs are slender with relatively long legs. They = range=20 from about 3/8 to =BD inch long and are tan to reddish-brown. The body = tapers=20 toward the head which is armed with a long beak. Damsel bug nymphs = resemble=20 adults but are smaller and lack wings.=20

In East Texas, damsel bugs usually are found at densities of less = than one=20 per 10 meters of row with occasional peaks of about two per meter = between=20 mid-June and mid-August. However, in some years and in some areas they = reach=20 densities of up to 10 per meter.=20

Damsel bugs feed on bollworm-budworm eggs and larvae as well as = aphids,=20 fleahoppers, Lygus bugs, leafhoppers and spider mites. They also prey on = other=20 predators such as minute pirate bugs and bigeyed bugs.=20

Assassin bugs (Zelus and = Sinea=20 spp.)=20

3D"Leafhopper The most abundant assassin bugs in cotton are the = leafhopper=20 assassin bug (Zelus renardii [Kolenati]) and the spined = assassin bug=20 (Sinea diadema [Fabricius]). The leafhopper assassin bug adults = are=20 about =BD inch long and of moderate to slender build. They are various = colors=20 including red, brown and yellowish green. The elongated narrow head is = armed=20 with a strong beak. The nymph resembles the adult but is smaller and = wingless.=20

The spined assassin bug also is about =BD inch long and is almost = uniformly=20 dark brown to dull reddish brown. Abdomen margins are expanded and flat = with a=20 pale spot on the rear margin of each segment. The front legs are covered = with=20 spines. Nymphs resemble the adults but are wingless and distinctly = sway-backed.=20

Assassin bugs peak at about one per 2 meters of row during July but = average=20 only seven per 100 meters during the season. Dispersal reaches a peak = during the=20 fall. They are predators of cotton fleahoppers and bollworm-budworm eggs = and=20 larvae and are one of the few predators that can readily capture and = consume an=20 adult boll weevil, as well as larger Heliothis larvae.=20

Spiders

Spiders are important in regulating pest insects but are not numerous = enough=20 to control excessive numbers that occur during major outbreaks. Along = with other=20 mortality agents, spiders often keep pests below densities that cause=20 unacceptable cotton losses.=20

Striped lynx (Oxyopes salticus = [Hentz])=20

3D"Striped The striped Lynx is a hunting spider and builds no web. = It can be=20 identified in the field by its habit of running rapidly and erratically = and=20 jumping. It is a brownish colored spider about 1/4 to 3/8 inch long with = four=20 longitudinal grey stripes behind the eyes on the front half (the = carapace). Two=20 black lines extend down the face and the jaws (chelicerae). Legs have = large=20 spines and the abdomen tapers to a point. Immatures resemble adults but = are=20 smaller.=20

This spider usually is the most abundant spider in East Texas cotton = fields,=20 averaging more than one per 2 meters of row with peaks of two per meter. = Numbers=20 of striped Lynx build throughout the growing season, peaking in = September. They=20 reproduce throughout the growing season so immature spiders are moving = into=20 cotton fields almost continuously. Immatures often are abundant on green = vegetation during the winter.=20

The striped Lynx preys on cotton fleahoppers, Lygus bugs and small=20 bollworm-budworm larvae.They do not readily prey on bollworm-budworm = eggs, but=20 in Arkansas they account for 11 to 14 percent of all arthropod predation = on=20 second-instar bollworm larvae. They may consume about one egg or small = larva=20 every 2 days.=20

Celer crab spider (Misumenops = celer=20 [Hentz])

3D"Celer The celer crab spider is also a hunting spider, does not = weave a=20 web and usually is found in plant terminals. The four front legs are = longer than=20 the four rear legs, and there usually are two longitudinal stripes = behind the=20 eyes on the carapace. Also, the rear half of the abdomen often has a "V" = shaped=20 mark pointing toward the rear. Adults reach a length of about 1/4 inch. = They are=20 yellow, green or white with reddish markings. Immature stages resemble = adults=20 but are smaller. These crab spiders enter the cotton field early but = reach a=20 peak density of about one per meter of row in October and November in = East=20 Texas. The seasonal average density is about one per 10 meters of row.=20

Celer crab spiders feed on cotton fleahoppers, Lygus bugs and a wide = range of=20 other insects, including bollworm-budworm eggs and worms; they may = consume about=20 one egg every 2 days and one worm per day.=20

Winter spider = (Chiracanthium=20 inclusum [Hentz])

3D"Winter The winter spider is found during the day in silken nests = under=20 leaves or bracts of squares. These hunting spiders do not spin a typical = web and=20 hunt primarily at night. The adult is about 1/3 inch long and cream to = pale=20 green with dark brown chelicerae. A faint, median longitudinal band is = found on=20 the front half of the abdomen. Young spiders resemble the adults.=20

Peak numbers of up to one per meter of row may be found during July = and=20 August. Average densities of about one per 10 meters are found = throughout the=20 season. Immatures float through the air on webs (ballooning) from March = to=20 December but most commonly from August through November.=20

Immatures of the winter spider feed on bollworm-budworm eggs, = possibly taking=20 14 per day when eggs are abundant. A second instar winter spider = consumes an=20 average of 3 =BD first instar bollworms in I =BD hours in the laboratory = and about=20 three per day in the field. They prefer bollworms and other species of = worms and=20 flies but also eat bugs, grasshoppers and beetles.=20

Star-bellied orb weaver = (Acanthepeira=20 stellata [Walckenaer])=20

3D"Star-bellied Large webs spun between cotton rows often are made by the = star-bellied orb weaver, frequently the most abundant web-spinner in = cotton.=20 When the web is found, the spider may be located under a leaf to which = one of=20 the foundation lines of the web has been=20

attached. One of its legs touches the line to alert the spider when = prey is=20 caught in the web.=20

Identification is relatively simple because of the characteristic = star-shape=20 of the abdomen formed by a series of cone-like bumps (protuberances). = The spider=20 is brown with a white spot on the anterior portion of the abdomen. = Adults reach=20 a length of about 3/8 inch. Adults and immatures differ only in size.=20

Numbers generally increase as the season progresses, reaching a peak = in=20 August and September when large numbers of immatures cover the = countryside.=20 Densities in cotton fields average about one per 10 meters of row but = peak at=20 one per 2 meters.=20

The star-bellied orb weaver does not hunt its prey but waits for prey = to get=20 caught in its web. Thus, it is likely that most of its prey fly or jump = into the=20 web. Adult fleahoppers, bollworm-budworm moths, bigeyed bugs, pirate = bugs,=20 damsel bugs and honeybees are frequent prey of this spider. There is no = evidence=20 that this spider feeds on bollworm or budworm eggs under field = conditions, but=20 it may consume one small larva every 2 days.=20

Grey dotted spider (Aysha = gracilis=20 [Hentz])=20

3D"Grey This spider is very similar in behavior and appearance to = the=20 winter spider. It is a night hunter and spends the day in a silken nest = under=20 leaves or bracts of squares. The adult is about 1/4 inch long and yellow = with a=20 darker anterior end and brown jaws (chelicerae). A pair of longitudinal = grey=20 bands is found on the back (carapace) between the eyes and the abdomen, = and=20 reddish-brown to black spots form two indistinct longitudinal bands on = the=20 abdomen. The legs are covered with spines; this helps distinguish it = from the=20 winter spider which has smaller and less distinct spines on the legs. = Immatures=20 resemble adults but markings are paler.=20

Peak numbers occur during August or September when one per 2 meters = of row=20 may be present. Usually fewer than one per 10 meters are found.=20

Little is known of its feeding habits in cotton, but it preys on = about one=20 bollworm-budworm egg every 10 days and one small worm every 2 days. = Otherwise=20 its prey is similar to that of the winter spider.=20

Black and white jumping spider = (Phidippus=20 audax [Hentz])=20

3D"Black Common around homes and gardens, this spider hunts in the = daytime=20 and often is seen on window screens. It is common in cotton. Its black = color is=20 contrasted with three white spots on the abdomen of adults. These spots = are=20 reddish-orange on the immatures which also have reddish legs. Adults may = reach =BD=20 inch in length and have large eyes on the front of the head. When = attacking,=20 they crawl to within striking distance and jump on their prey.=20

This spider generally increases its numbers throughout the season = reaching a=20 peak of about one per meter of row in August. Average density is about = one per=20 10 meters of row.=20

Immature spiders feed on bollworm-budworm eggs and may eat five per = day. This=20 spider can capture and eat all larval stages of the bollworm-budworm; = larger=20 larvae are taken by the larger spiders. Under field conditions they may = each eat=20 about one worm every 2 days. Even boll weevil adults and = bollworm-budworm moths=20 are attacked. Immature spiders capture and consume fleahoppers, Lygus = bugs and=20 pink bollworm adults.=20

Long jawed orb weaver = (Tetragnatha=20 laboriosa [Hentz])=20

3D"Long This elongated spider, less than 1/4 inch long, uses a = web placed=20 between branches of the cotton plant or between rows to catch its prey. = Males=20 have long jaws (chelicerae) from which this species gets its name. They = are=20 light yellow with a silvery abdomen. This spider often is found in the = center of=20 its web, which has an open hub.=20

A peak of about one per meter of row occurs in August with an average = density=20 of about one per 5 meters of row.=20

Feeding habits of this spider are not fully known, although they feed = mostly=20 on flying insects caught in their webs such as fleahoppers, leafhoppers = and=20 small moths.=20

Ridgefaced crab spider = (Misumenoides=20 formosipes [Walckenaer])=20

3D"Ridgefaced This crab spider is a hunter that usually waits in ambush = for its=20 prey in the terminal of the cotton plant. It can change color to match = its=20 background but is generally creamy-white to yellow or yellowish-brown = with=20 slightly darker sides. The abdomen may be unmarked or it may have red or = brown=20 markings that form a "V" pointing to the head. A white ridge crosses the = face=20 under the eyes.=20

An average density of only about one per 100 meters has been observed = in May,=20 although peaks of up to one per 10 meters have been observed. This = spider feeds=20 on fleahoppers and bollworm-budworm larvae and perhaps other insects.=20

Use of Predators

Current research in East Texas indicates that if any combination of = the 16=20 predators discussed in this publication is found in the cotton field in = numbers=20 qual to or greater than bollworm-tobacco budworm eggs, insecticides = usually are=20 not needed. For example, if an average of 10 eggs and 10 predators are = counted=20 per meter of row, no insecticide is needed. Use caution during rainy = weather=20 when the predation level may decrease. However, if the rains do not = continue=20 more than I or 2 days, the predators may recover and prevent = unacceptable=20 losses.=20

3D"Beat-bucket Careful sampling of these predators is as important as = sampling=20 pests. Thus, securing the services of a well-trained field scout or = consultant=20 is a good investment. Farmers can also easily learn to do this = themselves. A=20 "beat-bucket" sampler is recommended as a simple and efficient tool for = sampling=20 predators and some pests. To make one, cut the bottom 10 inches from a = white or=20 light-colored S-gallon plastic bucket. To use the sampler, place the top = 10=20 inches of the cotton plant inside the bucket and give five separate, = rapid=20 shakes from side to side. Then even the smallest predators can be = counted in the=20 bottom of the bucket.

INSECT PARASITISM OF COTTON PESTS

Allen Knutson and G.T. Bohmfalk=20

All animals and plants, including insects, have natural enemies that = attack=20 their various life stages. These natural enemies are what help keep = explosive=20 pest outbreaks from occurring. Important natural enemies of cotton pests = include=20 insect parasites.=20

A parasite is an organism that lives in or on another usually larger=20 organism. Parasites of insects, termed parasitoids, are a special group = of=20 parasites since they kill their host. An insect parasite requires only = one host=20 to develop to the adult stage, which is free living.=20

Insect parasites feed internally or externally on their hosts. An = external=20 larval parasite (Euplectrus hircinus [Say]) is seen = parasitizing a=20 bollworm larva. The host may become paralyzed before it dies from the = parasite's=20 feeding. Insect parasites may attack any life stage oftheir host, but = most=20 parasitize the egg or larval stage. Egg and larval parasites lay their = egg in a=20 host egg or larva. The parasite egg hatches into a larva that kills the = host egg=20 or larva. Some insect parasites lay an egg in the host egg, larval or = pupal=20 stage, and the parasite finally kills a later stage (larva, pupa or = adult).=20

3D"External 3D"Trichogramma

Tiny wasps called Trichogramma are typical insect parasites. = This=20 wasp parasitizes bollworm and budworm eggs. The adult female wasp = deposits her=20 egg inside the bollworm egg. The wasp's egg hatches into a larva which = feeds in=20 the bollworm's egg. The full grown Trichogramma larva = transforms into=20 the adult wasp which escapes from the dead bollworm egg. The female wasp = mates=20 and flies in search of other bollworm eggs to parasitize, thus repeating = the=20 life cycle. Trichogramma also parasitizes other moth eggs. 3D"Figure =20

In addition to Trichogramma wasps, bollworms and tobacco = budworm=20 larvae often areparasitized by other fly and wasp species. Many = insecticides=20 applied to cotton destroy insect parasites, which may result in economic = infestations of bollworms, budworms or other cotton pests. The boll = weevil, in=20 contrast, has relatively few effective natural enemies, which explains = in part=20 why it is such a devastating pest.=20

Most parasites are so small and secretive they are seldom seen and = their=20 benefit is not always evident. Other parasites, including tachinid = flies,=20 braconid wasps and ichneumonid wasps, are larger and more often seen. = Parasitic=20 insects are valuable allies in keeping cotton pest populations below = damaging=20 levels.=20

Recognizing the value of parasitic insects and preserving natural = enemies=20 whenever possible is an important component of cotton integrated pest=20 management.

THE COTTON PLANT

R.B. Metzer=20

The cotton plant usually is considered an annual, although it is a = perennial=20 in some parts of the world where it is grown commercially. The woody, = herbaceous=20 plant has a long tap root and attains a height of 2 to 5 feet or taller = under=20 favorable conditions. Cotton grows best with high temperatures and = adequate soil=20 moisture and fertility. Air temperatures in the 90=B0 to 95=B0 F. range = are=20 considered optimum for growth. Little or no growth can be expected at = 60=B0 F. or=20 below. Temperatures in excess of 100=B0 F. for several days can be = unfavorable,=20 especially if soil moisture becomes deficient. The tap root grows = downward into=20 moist soil at a rate of about 1 inch per day for 5 weeks or longer. = During the=20 growing season the daily growth rate of roots may average =BD inch.=20

When cotton emerges, the first leaf structures are called = cotyledonary or=20 seed leaves. They appear on the lowest node and are borne on opposite = sides of=20 the main stem. The nodes above the seed leaves bear a single true leaf. = These=20 leaves have a spiral arrangement around the stem. The true leaves have = five or=20 more clearly defined lobes. At the base of each main stem leaf, in the = angle=20 between the leaf and the stem, there are two and sometimes three buds. = They are=20 called axillary buds and give rise to the vegetative and fruiting = branches. The=20 vegetative branches normally are restricted to the lower nodes on the = stem. In=20 most American upland cottons the first fruiting branch begins developing = at the=20 fifth or sixth node above the seedling leaves. The fruiting branches = produce=20 floral buds, called cotton squares that develop into bolls. Flowers = (blooms) are=20 creamy white when first open. Fertilization occurs on day that flowers = open; it=20 turns pink the day after anthesis. Then boll development begins. The = interval=20 between corresponding nodes on successive fruiting branches (vertical = flowering=20 interval) is 2 to 3 days, and the interval between successive flowers on = the=20 same fruiting branch (horizontal fruiting interval) is 5 to 6 days.=20

Flowering usually begins about 7 to 11 weeks after planting. = Determinate=20 varieties stop growing after boll development. In contrast, = indeterminate=20 varieties usually are late-maturing and continue flowering until stopped = by=20 frost, drought, insect attack or some other cause. The flower bud or = square=20 usually is discernible 3 to 4 weeks before the flower opens. Many of the = buds,=20 squares, flowers or young bolls drop off naturally. This is called = shedding. As=20 a result, it is estimated that only 35 to 40 percent of the squares = produce=20 mature bolls under normal conditions. The most critical time for = developing=20 fruit is 3 to 10 days after pollination. The period between flowering = and=20 opening of mature bolls is 6 to 8 weeks, depending on growing = conditions. Cloudy=20 conditions and below normal temperatures during this period can increase = the=20 boll maturation period.=20

Fruit of the cotton plant is the enlarged 3- to 5-loculed ovary = commonly=20 referred to as a cotton boll. Mature bolls vary in size and shape = depending on=20 the variety and environmental conditions but usually are 1 1/2 to 2 = inches in=20 diameter. Bolls set during the first 3 weeks of fruiting usually are the = largest=20 and contain the highest quality fiber. Normally, late set bolls are = smaller and=20 may contain finer and less mature fiber, depending on temperature and = moisture=20 levels during the boll maturation period. Normally, 65 to 90 bolls are = required=20 to produce a pound of seed cotton. However, some varieties produce = relatively=20 small bolls that may require 100 bolls or more to produce a pound of = seed=20 cotton.=20

Fiber length development is determined the first 3 weeks after = flowering.=20 Increase in micronaire value as well as fiber strength occurs during the = next 3-=20 to 4-week period after length is established. Moisture stress during the = first 3=20 weeks after flowering can restrict fiber length. Adequate temperature is = a=20 critical factor in determining the strength and maturity of lint.=20

The cotton plant can adapt to different environmental stresses. In a = severe=20 drought, the plant may be only 6 inches high but still capable of = producing=20 several bolls. A plant of the same variety grown under adequate moisture = or=20 irrigation may be 5 feet in height and produce 40 to 50 bolls. Depending = on=20 other stresses, the number of bolls that develop is kept in balance with = the=20 vegetative growth of the plant. The fruiting-vegetative balance is = affected by=20 several factors. A shift to vegetative growth is associated with = excessive=20 nitrogen and soil moisture, coupled with cool, cloudy conditions. This = shift is=20 intensified even further if initial fruit set is poor or if excessive = shedding=20 has taken place. Warm, sunny weather and adequate but not excessive = moisture and=20 nitrogen and little insect stress encourage fruiting.=20

A population of 30,000 to 65,000 plants per acre is the optimum range = for=20 maximum yields. Although the cotton plant can adjust to variable = spacing,=20 uniform distribution of plants in the row produces the highest yields. A = uniform=20 plant spacing produces the highest yields. Under dryland conditions a = plant=20 population in the lower range is desirable, especially in low rainfall = areas.=20

Cottonseed are frequently planted at rates higher than necessary to = achieve=20 adequate stands. A dense stand results in crowding in the drill-row. = High plant=20 population encourages fruit set at a higher position on the plant, which = contributes to delayed maturity. Severe crowding can result in barren = plants=20 that function as weeds because they compete for moisture and nutrients. = Barren=20 plants usually are taller and are therefore more susceptible to lodging. =

The time required for specific growth and development stages in the = cotton=20 plant is influenced primarily by planting date, variety, available = moisture and=20 temperature during the fruiting period. Other factors such as soil type = and the=20 level of insect, weed and disease pressure also can alter the time = required to=20 reach various growth and fruiting stages. Some important growth and = fruiting=20 variables of the cotton plant are presented below:

**Average Growth and Fruiting of the Cotton=20 Plant**
Planting to emergence	4 to 10 days
Emergence to first true leaf	8 days
Emergence to second true leaf	9 days
Second true leaf to pinhead square (seventh node)	18 to 21 days
Pinhead square to matchhead square	9 to 10 days
Matchhead square to first one-third grown square	3 to 6 days
First one-third grown square to first white bloom	12 to 16 days
First white bloom to first open bolls	40 to 60 days
Harvest bolls set first 4 weeks of blooming	96%
Percent blooms that make bolls	35 to 40 %

Most fruit is set in the first 3 to 4 weeks of blooming. Squares are = more=20 easily shed than bolls. Once bolls are 12 days old or older, they = usually will=20 not shed unless the plant suffers severe stress (temperature, moisture, = insects=20 or disease).

CROP MONITORING (SCOUTING)

G.T. Bohmfalk=20

Integrated pest management evolved as several control techniques in = harmony=20 to keep pests below damaging levels. Experience shows that it is better = to=20 manage pests by using multiple suppression tactics rather than a single = one.=20 This includes using natural controls such as predators, parasites and = pathogens=20 and cultural strategies in combination with well timed insecticide = applications=20 in an integrated approach. Along with the development of this = multitactic=20 management system is an increased awareness of "in-field" insect and = plant=20 dynamics. This awareness is a result of constant crop monitoring of = pests and=20 beneficial insects and their effects on the cotton plant. This is the = basis of=20 cotton scouting.=20

Cotton scouting, or crop monitoring, is not a suppression tool itself = but=20 rather a means of gathering information on pests which will aid in = management=20 decisions. There is more to cotton monitoring than just checking for = insects. It=20 involves proper identification, a determination of pest and beneficial = insect=20 densities and their effects on the cotton crop as a whole. Insect = numbers=20 fluctuate a great deal. In cotton fields pests can increase to a point = where, if=20 not suppressed, they will cause economic damage to the crop. This pest = density=20 is called an economic threshold and is illustrated in figure 3. =

The economic threshold is determined by research and practical = experience. It=20 has been approximated for most cotton pests but may vary with the area = in which=20 the crop is grown. For current information on the economic thresholds of = cotton=20 pests in your area, contact an Extension entomologist or your county = Extension=20 agent or refer to the Management of Cotton Insects series = appropriate=20 for your region.=20

Since evaluating the economic threshold involves observing a pest = population=20 that is truly dynamic and rises and falls periodically, there must be = some=20 method of systematic sampling on a regular basis. Sampling in cotton = usually is=20 done once or twice a week. Sampling results should be recorded in an=20 easy-to-understand method (figure 4). Frequently, during peak insect = activity,=20 fields must be scouted or monitored more than twice a week. It is also = critical=20 to monitor the effects of other factors such as weather, disease, = herbicidal=20 injury, etc.

Insect populations often are concentrated in certain areas of a = field.=20 Insects are not distributed uniformly in the field and do not infest all = fields=20 in an area. Large areas of the field must be covered each time. Scouting = aids in=20 determining the kinds of insects present, their location, relative = abundance and=20 distribution in the field and numbers of insects or amount of insect = damage to=20 the plants. Scouting only near turn rows and field margins is inadequate = for=20 determining insect infestation. To cover the field completely, walk a = field in a=20 "zigzag" pattern crossing it diagonally from one corner to another or = walk it=20 in-a wide arc taking samples from several plants (figure 5). The pattern = followed should allow adequate sampling throughout the field, including = the=20 center, the sides and corners. Sampling pattern should be determined by = the=20 shape and size of the field. Avoid sampling in the same area of the = field on=20 successive sampling days.

The numbers and kinds of insects in a field usually are determined = with=20 reasonable reliability by examining squares and leaves from the terminal = or=20 entire plant or by sweeping the top part of a plant. Samples also may = include=20 any insects in the blooms, the rate at which squares are being set. the = blooming=20 rate. observable damage to bolls and the growth rate of plants. = Determine any=20 "natural" excessive square or boll shed when making an evaluation. = Remember to=20 consider the effect the weather has on the crop when evaluating fruit = loss.=20

The part of the plant examined in a sample varies with the insect = pest being=20 sampled. Many insect infestations are assessed by taking "square" = samples and=20 determining the percent damaged by the particular insect. Other insects = may be=20 found primarily on terminal buds, so the terminal may be the best = sampling unit.=20 Taking leaf samples and counting the number of insects present or the = amount of=20 leaf damage provide infestation information for many leaf-feeding = species.=20 Sample plant bugs and other insects by using an insect sweep net. Black = light=20 traps and pheromone traps are survey tools useful for = estimating the=20 relative density of some insects in the area.=20

Take samples or count samples from the bottom, middle and top = portions of the=20 plants. Taking all the samples from the top can give an inaccurate count = and let=20 insect infestations concentrated on the lower parts of the plants go = undetected.=20 This is especially true for bollworms and tobacco budworms and their = eggs, which=20 are sometimes laid lower on plants in the hotter part of the season.=20

The number of samples taken when scouting a field usually is dictated = by=20 field size and field variability. In fields of 40 acres or less, usually = only=20 two samples are made. A single sample should consist of examining at = least 25=20 individual plants at random or all plants in a predetermined length of = row (6.5=20 feet or 13 feet) at each site. Make a record of pests, beneficials and = plant=20 damage. The data from all samples are totaled or averaged for the field. = The=20 number of samples should increase as the field size increases. Fields = between 40=20 and 100 acres usually have four samples taken. In fields exceeding 100 = acres, a=20 sample is added for every additional 50 acres. For specific sampling=20 instructions refer to the Management of Cotton Insect series for your = region.=20

MONITORING COTTON FRUITING RATE

Monitoring the cotton fruiting rate is an excellent indicator of the = growth=20 and maturity of the cotton plant. Usually problem fields are detected = promptly=20 if fruiting rates are monitored accurately. Early fruiting is a = desirable=20 quality and frequent monitoring gives a good indication of crop set. To = monitor=20 fruiting rates, mark a point on a row and count 100 consecutive = one-third grown=20 (1/4 inch in diameter) or larger green squares. Record the number of row = feet=20 required to gain that count. One-third grown squares are preferred = feeding- and=20 egg-laying sites for boll weevils. This general square size also is = preferred by=20 bollworms/tobacco budworms; thus, it is a key indicator of potential = damage.=20 Also record the number of bolls observed in this distance. Later in the = season,=20 when squares are few, count 100 bolls, both green and open, and record = the=20 number of row feet required to make the count. To estimate the number of = squares=20 or bolls present per acre, divide the number of row feet required to = gain a=20 count of 100 squares or bolls into the number of row feet per acre = (13,068=20 running row feet for 40-inch rows and 13,756 running row feet for = 38-inch rows)=20 and multiply by 100.

Bolls or squares per acre =3D  13,068 (or =
13,756)     X 100
			   Number row feet recorded
			  for 100 consecutive bolls
				 or squares=20

MANAGEMENT DECISIONS

R.E. Frisbie and G.T. Bohmfalk=20

The decision to control cotton insects and mites with insecticides = should be=20 made with care. Use insecticides only if the economic threshold has been = reached, based on insect scouting results, the stage of plant growth and = rate of=20 fruit production. With the exception of a few microbial insecticides,=20 insecticide applications typically depress beneficial (predator and = parasite)=20 populations that are important natural control agents. Low rates of = certain=20 insecticides such as ovicides have been less damaging to beneficial = populations.=20 Insecticidal control of one pest often affects the beneficial population = suppressing another pest. The most frequent example is an insecticide = applied=20 near or at the bloom stage for boll weevil, which destroys beneficial = insects=20 that help keep the bollworm-tobacco budworm in check.=20

The growth stage of the cotton plant, in relation to its = vulnerability to=20 insect attack, is important to consider. One may view these = susceptibility=20 periods as "windows" in crop growth. Figure 6 illustrates these windows = and=20 their relationship, in time, with the plants growth stages. For example, = the=20 window for the cotton fleahopper is from pinhead size square through = one-third=20 grown square. After this period the cotton fleahopper usually does not = have the=20 potential to inflict damage. Windows of key pests may overlap and make = decisions=20 more difficult and risky. The best example of this is, again, with the = boll=20 weevil and the bollworm-tobacco budworm complex. The window for the boll = weevil=20 begins at one-third grown square and generally ends when a boll reaches = 10 to 12=20 days of age. The window for the bollworm-tobacco budworm begins around = the=20 one-third grown square stage and extends past and overlaps with the boll = weevil=20 window.

3D"Figure =20

Research shows that the most critical period of fruit formation is in = the=20 first 30 to 40 days of blooming. The highest quality and quantity of = lint is=20 produced at this time. Thus, damage inflicted in this window can be most = costly.=20 Strategies in cotton insect management show that the boll weevil can be = managed=20 effectively in various parts of the state by altering the planting date, = if=20 practiced uniformly, or by selected insecticide applications early in = the boll=20 weevil window (first one-third grown square stage), based on field = scouting=20 information. Either of these strategies, if properly practiced in most = years,=20 neutralizes the boll weevil as a pest, allows theplant to set and hold = early=20 fruit and delays boll weevil build-up until the critical 30 to 40 days = of=20 blooming can take place. Of equal importance, this strategy has little = effect on=20 beneficials that naturally control bollworm-tobacco budworm populations. =

By viewing growth of the cotton plant as a series of susceptibility = windows,=20 management decisions and scouting information can be targeted to deal = with the=20 specific major pests attacking cotton. By considering the pest within a = certain=20 window (time-frame), its damage can be assessed relative to the growth = stage and=20 its potential to destroy harvestable fruit.

OTHER INSECT PEST MANAGEMENT PRACTICES

A field sterile of all pest insects is neither necessary nor = desirable.=20 However, do not allow harmful insects to reach numbers that will cause=20 unacceptable losses. Keep them below the economic threshold. Excessive = use of=20 nitrogen fertilizer causes cotton plants to stay green and stimulates = rank=20 growth. The excessive, rank growth can reduce cotton yield and increase = damage=20 from the bollworm-tobacco budworm. Rank growth also makes it difficult = to=20 achieve thorough plant coverage with insecticides.=20

Cotton planted too thick becomes tall and "leggy" before it sets = squares.=20 This condition is often wrongly blamed on cotton fleahoppers. Maximum = yields are=20 produced when cotton is planted at a rate of two to four plants per foot = of row=20 on 40-inch rows. Current research in this area on narrow row spacings is = being=20 evaluated for use by cotton producers in various production areas.=20

Foliage and fruit on cotton stalks remaining in a field after harvest = can=20 supply a food source for boll weevils, bollworms, tobacco budworms and = pink=20 bollworms that allows these pests to continue growing and reproducing. = This=20 increases the number of overwintering pests. Cut stalks and plow fields=20 immediately after the cotton is picked or stripped.=20

By planting cotton at a uniform time throughout an area, many = problems with=20 boll weevils, bollworms and tobacco budworms and pink bollworms can be = reduced=20 greatly. These insect pests are attracted to the older cotton plants of = early=20 planted cotton. Late season damage by bollworm-tobacco budworm often is = greater=20 on later planted cotton which remains green and lush longer than earlier = planted=20 cotton. Later planted cotton also may have more squares which aid in the = development of high numbers of boll weevils.

INSECT IDENTIFICATION AND DAMAGE-DIAGNOSTIC KEY

G.T. Bohmfalk=20

This simple key is designed to help the producer diagnose damage = caused by=20 insects and spider mites. Although it is pest-oriented, cues are = provided to aid=20 in separating pest identification from beneficial identification. = Extension=20 entomologists can assist with identifications .=20

Symptom	Suspected insect
Seeds
Seeds eaten, resulting in poor stand	Fire ant or seed-gathering ant (not common)
Seeds hollowed out	Wireworms (not common)
Seedling
Stems cut off evenly at or near ground level; chewed at = base	Cutworms Grasshoppers
Cotyledons and first rue leaves deformed or "cupped downward" = with=20 honeydew present	Aphids or whiteflies (Whiteflies may not deform leaves but = honeydew is=20 present.)
Cotyledons and first true leaves deformed, "cupped upward" with = no=20 honey dew but with a silvered appearance	Thrips
Ragged holes eaten in leaves	Armyworms Cabbage=20 looper (grasshoppers)

**Established Stand Through=20 Squaring**
Symptom	Suspected Insect
Leaves
Discolored, reddened or "mottled" some slight webbing obvious = on=20 underside	Spider mite
With "shot-hole" appearance; mining and horseshoe-shaped = larvae=20 present	Cotton leaf perforator
Skeletonized with numerous small larvae present	Armyworms
Numerous large holes and feeding on margin	Cabbage looper or grasshoper
Ragged and completely eaten with "woolybear" caterpillars=20 present	Saltmarsh caterpillar
Rolled with webbing present	Omnivorous leaf roller
Squares Blackened or "blasted"	Lygus bugs Cotton fleahopper
Eaten into with frass present, hole circular and square may = be=20 hollowed out	Bollworm and/or tobacco budworm
Eaten into with clean feeding site hole slightly flattened = and=20 square not completely hollowed out	Cotton square borer
Bracts primarily fed on	Beet armyworm
Tiny feeding sites with bright orange to yellow excrement = present;=20 small wart-like, egg-laying sites	Boll weevil
Blooms
Holes in base	Armyworms Bollworm and/or tobacco budworm
Rosetted petals	Pink bollworm
Disfigured, twisted and coarse	Lygus (plant) bug, cotton fleahopper or stink = bug
Terminal Tunneling in terminal	Bollworm-budworm, fall armyworm or omnivorous leaf=20 roller

**Boll Set and Maturity**
Symptom	Suspected Insect
Bolls
Holes with frass present; hole circular	Bollworm and/or tobacco budworm Beet armyworm Fall=20 armyworm
Tiny holes with some yellow to bright orange frass and = wart-like,=20 egg laying sites	Boll weevil
Small depressions and "cat-facing"	Lygus bug Stink bug
Holes through boll, carpel walls and seeds	Pink bollworm
Soft, dark area with cream colored grub inside boll	Boll weevil

**Insect Egg Diagnosis**
Symptom	Suspect Insect
Occurring = singly
White to tan, eventually turning brown, spherical with lines = from=20 base to tip, usually in terminal areas or underside of = tender=20 young leaves and on stems	Bollworm and/or tobacco budworm
Slightly flat, greenish-white, laid on leaf surface=20 Blue-green, flat and laid singly over leaves	Cabbage looper
White or green supported by a silken stalk	Lacewing
White, oblong, lays on side on leaves and stems	Bigeyed bugs
Silken bag attached to plant	Spiders
White, oval, flattened, somewhat donut- shaped, surface = covered=20 with net-like lines	Cotton square borer

Occurring in = groups
Pale green covered with scales and usually deposited on = underside=20 of leaves	Armyworms
Blue-green, flat and overlapping like shingles usually on=20 upperside of leaves	Omnivorous leaf roller
Brownish, "barrel-shaped," tightly packed	Stink bugs
Orange, shiny and cigar-shaped with few in cluster	Lady beetle

**Insect Larval Diagnosis**
Symptom	Suspect Insect
Found outside of bolls or squares =
Large, covered with dense yellow to brown hair	Saltmarsh caterpillar
Body with no hair, head capsule smaller than distal end, = pale=20 green, moves with looping action	Cabbage looper
Body having sparse hairs or spines; may be striped with = mottled=20 head capsule	Bollworm and/or tobacco budworm
Green with a tiny black spot above second true leg; without = even=20 small hairs	Beet armyworm
Black to dark green with yellow and brown stripes	Yellowstriped armyworm
Brown with dark patches on the sides; plae to white inverted = "Y"=20 on face of head capsule; body sparsely covered with short, = dark=20 hairs	Fall armyworm
"Football-shaped" with dense, velvet hair covering, bright = green=20 to yellow	Cotton square borer
Smooth, olive green with white spots on each segment; = darkened=20 head; slender worm often found rolled in cotton leaves	Omnivorous leaf roller
Tiny, white to dark green, marked with darkened areas, = foundwithin=20 the leaf itself or on the surface	Cotton leaf- perforator
"C"-shaped, variable in color, in or on the soil surface;=20 "C"-shaped when disturbed	Cutworms

Found only inside bolls or = squares
Small whitish or pink larva inside maturing boll	Pink bollworm
Small, grub-like larvar found in squares or developing = bolls	Boll weevil

ADDITIONAL BENEFICIAL INSECTS

=20

The information given herein is for educational purposes only. = Reference to=20 commercial products or trade names is made with the understanding that = no=20 dicrimination is intended and no endorsement by the Cooperative = Extension=20 Service is implied.

Educational programs conducted by the = Texas=20 Agricultural Extension Service serve people of all ages regardless of=20 socioeconomic level, race, color, sex, religion, handicap or national = origin.=20

Issued in furtherance of Cooperative Extension Work in = Agriculture=20 and Home Economics, Acts of Congress of May 8, 1914, as amended, and = June 30,=20 1914, in cooperation with the United States Department of Agriculture. = Zerle L.=20 Carpenter, Director, Texas Agr icultural Extension Service. the Texas = A&M=20 University System.
20M--7-82, Revision=20
ENT

Return to = top of=20 page

Return to=20 Departmental Home Page

Last modified: December 9, 1996 by Edgar Cross
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IDENTIFICATION, BIOLOGY AND SAMPLING OF COTTON INSECTS

TABLE OF CONTENTS

INTRODUCTION

IDENTIFICATION, BIOLOGY, EVIDENCE OFINFESTATION AND NATURE OF=20 DAMAGE

OCCASIONAL PESTS

PREDACEOUS INSECTS AND SPIDERS

Insects

Spiders

Use of Predators

INSECT PARASITISM OF COTTON PESTS

THE COTTON PLANT

CROP MONITORING (SCOUTING)

MONITORING COTTON FRUITING RATE

MANAGEMENT DECISIONS

OTHER INSECT PEST MANAGEMENT PRACTICES

INSECT IDENTIFICATION AND DAMAGE-DIAGNOSTIC KEY

ADDITIONAL BENEFICIAL INSECTS

IDENTIFICATION, BIOLOGY, EVIDENCE OF
INFESTATION AND NATURE OF=20 DAMAGE